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A mixture contains three similarly sized peptides P, Q and R. The peptide P is positively charged, Q is weakly negative and R is strongly negative. If this mixture is passed through an ion-exchange chromatography column containing an anionic resin, their order of elution will be

  1. P, Q, R
  2. R, Q, P
  3. Q, R, P
  4. P, Q and R elute together
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