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Phylum proteobacteria is subdivided into $\alpha-, \beta-, \gamma-, \delta-$ and $\epsilon-$ proteobacteria based on

  1. $G+C$ content
  2. $23S\; rRNA$ sequences
  3. $tRNA$ sequences
  4. $16S\; rRNA$ sequences
in Recombinant DNA technology and Other Tools in Biotechnology edited by
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