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Which of the following factors can influence the lag phase of a microbial culture in a batch fermentor?

  1. Inoculum size
  2. Inoculum age
  3. Medium composition
  1. $P$ and $Q$ only
  2. $Q$ and $R$ only
  3. $P$ and $R$ only
  4. $P, Q$ and $R$
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