An analysis of DNA-protein interactions was carried out using all DNA-protein complexes in the protein data bank (PDB). The frequency distribution of four amino acid residues, represented as P, Q, R and S, occurring in non-covalent interactions between the protein and DNA backbone is shown below.

Which one of the following is correct?
- P-Lys, Q-Arg, R-Gln, S-Glu
- P-Gln, Q-Glu, R-Lys, S-Arg
- P-Asn, Q-Asp, R-Arg, S-Lys
- P-His, Q-Glu, R-Gln, S-Lys