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ATP biosynthesis takes place utilizing the $H^+$ gradient in mitochondria and chloroplasts. Identify the correct sites of $H^+$ gradient formation.

  1. Across the outer membrane of mitochondria and across the inner membrane of chloroplast
  2. Across the inner membrane of mitochondria and across the thylakoid membrane of chloroplast
  3. Within the matrix of mitochondria and across the inner membrane of chloroplast
  4. Within the matrix of mitochondria and Within the stroma of chloroplast

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