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​​​​Which one of the following is true for piRNAs?

  1. piRNAs silence transposable elements in germ cells
  2. piRNA is the abbreviation of $P$-element interacting RNA
  3. piRNAs modify the $2^{\prime}-\mathrm{OH}$ of ribose with methyl group
  4. piRNA is a long non-coding RNA

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